It is the thirty-second millennium and the Imperium is at peace. The Traitor Legions of Chaos are but a distant memory and the many alien races that have long plagued mankind are held in check by the Space Marines. But an ork invasion of unparalleled scale throws the Imperium into disarray. As worlds burn and renowned Space Marine Chapters are destroyed, can the Imperium pull together to defeat the Beast?

The primarchs, nigh-immortal sons of the Emperor of Mankind, take the stage in a series of short stories showcasing some of their great deeds and hideous acts from across the Great Crusade and the Horus Heresy.

The Night Lords were once among the most potent forces of the Imperium, Space Marines who used fear itself as their weapon. Now, cast adrift from the Emperor's light and hunted as heretics after their monstrous betrayal, the Night Lords clad themselves in symbols of death and fight the Long War, bringing pain and terror to all who worship the corpse-god of Terra. A summons from Warmaster Abaddon sends these rebels on a dangerous journey that leads inexorably to a conflict with the Emperor's chosen warriors, the Blood Angels.

A great darkness has befallen the galaxy, and the armies of Chaos are rampant. To survive, humanity must retaliate and take back what they have lost. By the will of the reborn primarch, Roboute Guilliman, is the Indomitus Crusade launched - a military undertaking that eclipses all others in known history. From the Throneworld of Terra does the Avenging Son hurl his fleets, their mission the very salvation of mankind.

On the grim death world of Fenris, the Space Wolves Chapter selects its aspirants from the best and brightest of the young tribesmen. Ragnar of the Thunderfist tribe finds his life changed forever when he is chosen. After being revived from a savage death on the battlefield, Ragnar is recruited into the fearsome Space Wolves Chapter. He is then thrown into a galactic war against the dark forces of Chaos. However, the implanting of the Canis Helix unleashes his primal instincts and Ragnar must fight to control the beast within him.

After seven long years of conflict, the Horus Heresy nears its end. But now, as the Warmaster Horus marches on Terra to rip his father from the Throne, the seat of the arch-traitor’s power lies empty. Cthonia, a world once dominated by brutal murder-gangs, is occupied by the Emperor’s praetorians—the indomitable Imperial Fists. For the Sons of Horus, this outrage demands a reckoning.

For a thousand years, the Sabbat Worlds have been lost to the Imperium, claimed by the dread powers of Chaos. Now, a mighty crusade seeks to return the sector to Imperial rule. And at the forefront of that crusade are Colonel-Commissar Ibram Gaunt and the Tanith First and Only - better known as Gaunt’s Ghosts. Trapped in the grinding trench warfare of Fortis Binary, the Ghosts find themselves drawn into a conspiracy to assassinate the crusade’s leader, Warmaster Macaroth.

After seven years of bitter war, the end has come at last for the conflict known infamously as the Horus Heresy. Terra now lies within the Warmaster’s sights, the Throneworld and the seat of his father’s rule. Horus’ desire is nothing less than the death of the Emperor of Mankind and the utter subjugation of the Imperium. He has become the ascendant vessel of Chaos and amassed a terrible army with which to enact his will and vengeance. But the way to the Throne will be hard as the primarch Rogal Dorn, the Praetorian and protector of Terra, marshals the defences.

Attached to Crusade Fleet Quintus - dubbed the Cursed Fleet by many - the Ultramarines of the Ithraca’s Vengeance are drawn to a stricken world. With millions enslaved, a malign necron technology siphons the souls of the innocent and heralds the Silent Kingdom’s expansion. The Ultramarines face an impossible decision: mount a desperate last stand to destroy the Pariah Nexus, or break away and damn the entire sector to bring word of this ancient foe’s resurgence to the only being capable of halting it - the Lord Primarch Roboute Guilliman.

When the world of Damnos is hit by cataclysmic earthquakes, an ancient force is awakened. Deep beneath the earth, the necrons rise from their slumber to decimate the human populace. All appears lost until salvation comes from the heavens – the Ultramarines brave an orbital bombardment to deploy their forces on Damnos, led by two legendary warriors – Captain Cato Sicarius and Chief Librarian Tigurius.